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2021 Mulder, T., Wilkins, L., Mortimer, B. and Vollrath, F. - Dynamic environments do not appear to constrain spider web building behaviour. The Science of Nature/Naturwissenschaften ; In Press

2020 Vollrath, F. and Krink, T. - Spider webs inspiring soft robotics. Journal of the Royal Society Interface e20200569 ; Open access Associated apps available here.

2020 Mulder, T., Mortimer, B. and Vollrath, F. - Functional flexibility in a spider’s orb web. Journal of Experimental Biology 223 Open access


2019 Frydrych, M., Greenhalgh, A. and Vollrath, F. - Artificial spinning of natural silk threads. Scientific Reports 9:(1) Open access

2019 Fu, C., Wang, Y., Guan, J., Chen, X., Vollrath, F. and Shao, Z. - Cryogenic toughness of natural silk and a proposed structure–function relationship. Materials Chemistry 3:(11) Open access

2019 Mi, R., Shao, Z. and Vollrath, F. - Creating artificial Rhino Horns from Horse Hair. Scientific Reports 9:(1) Open access

2019 Mortimer, B. - A Spider’s Vibration Landscape: Adaptations to Promote Vibrational Information Transfer in Orb Webs. Integrative and Comparative Biology 59:(6) access


2018 Holland, C., Hawkins, N., Frydrych, M., Laity, P., Porter, D. & Vollrath, F. - Differential Scanning Calorimetry of Native Silk Feedstock. Macromol Biosci. Open access

2018 Mortimer, B., Soler, A., Siviour, C.R., & Vollrath, F. - Remote monitoring of vibrational information in spider webs. The Science of Nature 105: (37) access

2018 Woods, A.K., Broomfield, J., Monk, P., Vollrath, F. & Glyn-Jones, S. - Dynamic shear analysis: a novel method to determine mechanical integrity of normal and torn human acetabular labra Bone Joint Res. 7: 440–446 access

2018 Hey Tow, K., Chow, D., Vollrath, F., Dicaire, I., Gheysens, T. & Thévenaz, L. - Exploring the Use of Native Spider Silk as an Optical Fiber for Chemical Sensing. J. Lightwave Technol. 36: (4) 1138-1144 access

2018 Shimanovich, U., Pinotsi, D., Shimanovich, K., Yu, N., Bolisetty, S., Adamcik, J., Mezzenga, R., Charmet, J., Vollrath, F. & Gazit, E. et al - Biophotonics of Native Silk Fibrils. Macromolecular bioscience 18: (4) access

2018 Wang, J., Guan, J., Hawkins, N. & Vollrath, F. - Analysing the structure and glass transition behaviour of silks for archaeology and conservation. Journal of the Royal Society, Interface 15: (139) access

2018 Vollrath, F., Mi, R. & Shah, D.U. - Ivory as an Important Model Bio-composite Curator: The Museum Journal 61 (1): 95-110 access


2017 Shimanovich, U., Ruggeri, S.F., De Genst, E., Adamcik, J., Barros, T.P., Porter, D., Muller, T., Mezzenga, R., Dobson, C.M., Vollrath, F., Holland, C. & Knowles, T.P.J. - Silk micrococoons for protein stabilisation and molecular encapsulation Nature Comms, 8: 15902 access

2017 Cotton, L.J., Vollrath, F., Brasier, M.D. & Dicko, C. - Chemical relationships of ambers using attenuated total reflectance Fourier transform infrared spectroscopy Geo. Soc. London, Special Publications, 448 access

2017 Koebley, S.R., Vollrath, F. & Schniepp, H.C. - Toughness-enhancing metastructure in the recluse spider's looped ribbon silk. Mater. Horiz., 2017, 4: 377-382 access

2017 Woodcock, J.W., Beams, R., Davis, S.D., Chen, N., Stranick, S.J., Shah, D.U., Vollrath, F. & Gilman, J.W. - Observation of Interfacial Damage in a Silk-Epoxy Composite, Using a Simple Mechanoresponsive Fluorescent Probe Advanced Materials Interfaces 4:10 access

2017 Elettro, H., Neukirch, S., Vollrath, F. & Antkowiak, A. - Drop-on-coilable-fibre systems exhibit negative stiffness events and transitions in coiling morphology. Soft Matter access

2017 Guan, J., Zhu, W., Liu, B., Yang, K., Vollrath, F. & Xu, J. - Comparing the microstructure and mechanical properties of Bombyx mori and Antheraea pernyi cocoon composites Acta Biomaterialia, 47: 60-70 access


2016 Monks, JN., Yan, B., Hawkins, N., Vollrath, F. & Wang, Z. - Spider Silk: Mother Nature’s Bio-Superlens Nano Lett, 16: 5842–5845 access

2016 Smitthipong, W., Suethao, S., Shah, D. & Vollrath, F. - Interesting green elastomeric composites: Silk textile reinforced natural rubber Polymer Testing, 55: 17–24 access

2016 Kraznov, I., Seydel, T., Greving, I., Blankenburg, M., Müller, M. & Vollrath, F. - Strain-dependent fractional molecular diffusion in humid spider silk fibres, J. R. Soc. Interface, 13 (122) access

2016 Hey Tow, K., Chow, D., Vollrath, F., Dicaire, I., Gheysens, T. & Thévenaz, L. - From spider webs to a fibre-optic chemical sensor EPFL Conference paper access

2016 Hey Tow, K., Chow, D., Vollrath, F., Dicaire, I., Gheysens, T. & Thévenaz, L. - Weaving our way towards a new generation of fibre-optic chemical sensors based on spider silk EPFL Conference paper access

2016 Hey Tow, K., Chow, D., Vollrath, F., Dicaire, I., Gheysens, T. & Thévenaz, L. - Towards a novel "silky" way to detect chemical species using spider threads EPFL Conference paper access

2016 Offord, C., Vollrath, F. & Holland, C. - Environmental effects on the construction and physical properties of Bombyx mori cocoons Journal of Materials Science51 (24): 10863–10872 access

2016 Dicko, C., Kasoju, N., Hawkins, N., & Vollrath, F. - Differential scanning fluorimetry illuminates silk feedstock stability and processability, Soft Matter, 12: 255-262 access

2016 Boulet-Audet, M., Holland, C., Gheysens, T. & Vollrath, F. - Dry-Spun Silk Produces Native-Like Fibroin Solutions Biomacromolecules, 17 (10): 3198–3204 access

2016 Vollrath, F. - The complexity of silk under the spotlight of synthetic biology Biochemical Society Transactions, 44 (4): 1151-1157 access;

2016 Kasoju, N., Hawkins, N., Pop-Georgievski, O., Kubies, D. & Vollrath, F. - Silk fibroin gelation via non-solvent induced phase separation Biomater. Sci., 4: 460-473 access

2016 Mortimer, B., Soler, A., Siviour, C. R., Zaera, R. and Vollrath, F. - Tuning the instrument: sonic properties in the spider’s web, J. R. Soc. Interface, 13 (22) access

2016 Guan, J., Wang, Y., Mortimer, B., Holland, C., Shao, Z., Porter, D. and Vollrath, F. - Glass transitions in native silk fibers studied by DMTA. Soft Matter, 12, pp. 5926-5936 access

2016 Elettro, H., Neukirch, S., Vollrath F. & Antkowiak, A. - In-drop capillary spooling of spider capture thread inspires hybrid fibers with mixed solid–liquid mechanical properties. PNAS, 113 (22): 6143-47 access


2015 Mortimer, B. & Holland, C. - The use of spider silk as a biomaterial. In: Basu, A. (ed.) Advances in Silk Science and Technology. Cambridge: Woodhead publishing

2015 Mortimer, B. & Vollrath, F. - Diversity and properties of key spider silks, Research & Knowledge, 1 (1): 32-42 access

2015 Mortimer, B. - Animal silks: denatured super proteins, Interface column in Biological Sciences Review, November issue.

2015 Mortimer, B., Holland, C., Windmill, J. F. C. & Vollrath, F. - Unpicking the signal thread of the sector web spider Zygiella x-notata. J. R. Soc. Interface, 12: 113 access

2015 Dicko, C., Kasoju, N., Hawkins, N. & Vollrath, F. - Differential scanning fluorimetry illuminates silk feedstock stability and processability. Soft Matter, advance access

2015 Boulet-Audet M., Vollrath F., & Holland C. - Identification and classification of silks using infrared spectroscopy. J. Exp. Biol., 218: 3138-3149 access

2015 Vollrath, F. - Spider Silk Encyclopedia of Nanotechnology, 1-11 access

2015 Astudillo, M., Thalwitz, G. & Vollrath, F. - Modern analysis of an ancient integrated farming arrangement: life cycle assessment of a mulberry dyke and pond system The International Journal of Life Cycle Assessment, 20 (10): 1387–1398 access

2015 Shah, D., Vollrath, F. & Porter, D. - Silk cocoons as natural macro-balloon fillers in novel polyurethane-based syntactic foams Polymer, 56: 93–101 access

2015 Elettro H., Vollrath F., Antkowiak A.,& Neukirch S., - Coiling of an elastic beam inside a disk: A model for spider-capture silk. International Journal of Non-Linear Mechanics, 75: 59-66 access

2015 Elettro H., Neukirch S., Antkowiak A., & Vollrath F., - Adhesion of dry and wet electrostatic capture silk of uloborid spider. Sci. Nat., 102: 41 access

2015 Koebley S.R., Thorpe D., Pang P., Chrisochoides P., Greving I., Fritz Vollrath F., & Schniepp, H.C., - Silk reconstitution disrupts fibroin self-assembly. Biomacromolecules, 16 (9): 2796-2804 access

2015 Mortimer, B., Guan, J., Holland, C., Porter, D., & Vollrath, F. - Linking naturally and unnaturally spun silks through the forced reeling of Bombyx mori. Acta Biomaterialia, 11: 247-255 access paper and audio slides

2015 Kronenberger, K. & Vollrath, F. - Spiders spinning electrically charged nano-fibres. Biology Letters, 11: 20140813 access


2014 Shah, D.U., Porter, D. & Vollrath, F. - Silk cocoons as natural macro-balloon fillers in novel polyurethane-based syntactic foams. Polymer. access

2014 Shah, D.U., Porter, D. & Vollrath, F. - Can silk become an effective reinforcing fibre? A property comparison with flax and glass reinforced composites. Composites Science and Technology, 101: 173-183. access

2014 Vollrath, F., Hawkins, N., Porter, D., Holland, C. & Boulet-Audet, M. - Differential Scanning Fluorimetry provides high throughput data on silk protein transitions. Scientific Reports, 4: 5625. access

2014 Shah, D.U., Vollrath, F., Porter, D., Stires, J. & Deheyn D.D. - Housing tubes from the marine worm, Chaetopterus sp.: biomaterials with exceptionally broad thermo-mechanical properties. Journal of Royal Society Interface, 11(98): 20140525. access

2014 Astudillo, M.F., Thalwitz, G. & Vollrath, F. - Life cycle assessment of Indian silk. Journal of Cleaner Production, 81: 158-167. access

2014 Mortimer, B., Gordon, S.D., Holland, C., Siviour, C.R., Vollrath, F., & Windmill, J.F. - The speed of sound in silk: Linking material performance to biological function. Advanced Materials, 26 (30): 5179-5183 access

2014 Shah, D.U. - Natural fibre composites: Comprehensive Ashby-type materials selection charts. Materials and Design, 62: 21-31. access

2014 Shah, D.U., Vollrath, F. & Porter, D. - Opportunities for silk textiles in reinforced biocomposites: Studying through-thickness compaction behaviour. Composites Part A: Applied Science and Manufacturing, 62: 1-10. access

2014 Holland, C., Vollrath, F. & Gill, H. - Horses and cows might teach us about human knees. Naturwissenschaften 101 (4): 351-354 access

2014 Boulet-Audet, M., Terry, A., Vollrath, F. & Holland, C. - Silk protein aggregation kinetics revealed by Rheo-IR. Acta Biomaterialia, 10 (2): 776-784 access


2013 Vollrath, F. & Edmonds, D. - Consequences of Electrical Conductivity in an Orb Spider’s Capture Web. Naturwissenschaften, 100 (12): 1163-1169 access

2013 Bauer, G., Friedrich, C., Gillig, C., Vollrath, F., Speck, T. & Holland, C. - Investigating the rheological properties of native plant latex. J. R. Soc. Interface, 11: 20130847 access

2013 Guan, J., Vollrath, F. & Porter, D. - Silk quality revealed using Dynamic Mechanical Thermal Analysis (DMTA). in '6th BACSA International Conference: Building Value Chains in Sericulture'. Padua, Italy. access

2013 Vollrath, F., Carter, R., Rajesh G.K., Thalwitz, G. & Astudillo, M.F. - Life Cycle Analysis of Cumulative Energy Demand on Sericulture in Karnataka, India. in '6th BACSA International Conference: Building Value Chains in Sericulture'. Padua, Italy. access

2013 Chen, F., Hesselberg, T., Porter, D. & Vollrath, F. - The impact behaviour of silk cocoons Journal of Experimental Biology, 216: 2648-2657 access

2013 Wang, Y., Porter, D. & Shao, Z. - Using Solvents with Different Molecular Sizes to Investigate the Structure of Antheraea Pernyi Silk. Biomacromolecules, 14 (11): 3936-3942 access

2013 Guan, J., Porter, P. & Vollrath, F. - Spider silk morphology for responsive materials Materials Research Society, 1498: 197-202 access

2013 Mortimer, B., Holland, C. & Vollrath, F. - Forced Reeling of Bombyx mori Silk: Separating Behavior and Processing Conditions. Biomacromolecules, 14 (10): 3653-3659 access

2013 Schniepp, H.C., Koebley, S.R. & Vollrath, F. - Brown Recluse Spider’s Nanometer Scale Ribbons of Stiff Extensible Silk. Advanced Materials, 25 (48): 7028-7032 access

2013 Davies, G.J.G., Knight, D.P. & Vollrath, F. - Chitin in the Silk Gland Ducts of the Spider Nephila edulis and the Silkworm Bombyx mori. PLOS ONE, 8 (8): e73225 access

2013 Shah, D.U. - Developing plant fibre composites for structural applications by optimising composite parameters: a critical review. Journal of Materials Science, 48 (18): 6083-6107 access

2013 Davies, G.J.G., Knight, D.P. & Vollrath, F. - Structure and function of the major ampullate spinning duct of the golden orb weaver, Nephila edulis. Tissue and Cell, 45 (5): 306-311 access

2013 Guan, J., Porter, D. & Vollrath, F. - Thermally Induced Changes in Dynamic Mechanical Properties of Native Silks. Biomacromolecules, 14 (3): 930-937 access

2013 Mortimer, B., Drodge, D.R., Dragnevski, K.I., Siviour, C.R. & Holland, C. - In situ tensile tests of single silk fibres in an environmental scanning electron microscope (ESEM). Journal of Materials Science, 48 (14): 5055-5062 access

2013 Horrocks, N.P.C., Vollrath, F. & Dicko, C. - The silkmoth cocoon as humidity trap and waterproof barrier. Comp Biochem Physiol A Mol Integr Physiol., 164 (4): 645-652 access

2013 Vollrath, F., Porter, D. & Holland, C. - The Science Of Silks. MRS Bulletin, 38 (1): 73-80 access

2013 Porter, D. & Vollrath, F. - Water mediated proton hopping empowers proteins. Softmatter, 9 (3): 643-646 access

2013 Porter, D., Guan, J. & Vollrath, F. - Spider Silk: super material or thin fibre? Advanced Materials, 25 (9): 1275-1279 access


2012 Yang, Y., Dicko, C., Bain, C.D., Gong, Z., Jacobs, R.M.J., Shao, Z., Terry, A.E. & Vollrath, F. - Behavior of silk protein at the air–water interface. Soft Matter, 8 (37): 9705-9712 access

2012 Hesselberg, T. & Vollrath, F. - The mechanical properties of the non-sticky spiral in Nephila orb webs (Araneae, Nephilidae). J. Exp. Biol., 215 (19): 3362-3369 access

2012 Drodge, D.R., Mortimer, B., Holland, C. & Siviour, C.R. - Ballistic Impact to Access the High-Rate Behaviour of Individual Silk Fibres. Journal of the Mechanics and Physics of Solids, 60 (10): 1710-1721 access

2012 Chen, F., Porter, D. & Vollrath, F. - Structure and physical properties of silkworm cocoons. Journal of the Royal Society Interface, 9 (74): 2299-2308 access

2012 Porter, D. & Vollrath, F. - Water mobility, denaturation and the glass transition in proteins. Biochimica et Biophysica Acta (BBA) - Proteins & Proteomics, 1824 (6): 785–791 access

2012 Wang, T., Porter, D. & Shao, Z. - The Intrinsic Ability of Silk Fibroin to Direct the Formation of Diverse Aragonite Aggregates. Advanced Functional Materials, 22 (2): 435-441 access

2012 Teshome, A., Vollrath, F., Rainaa, S., Kabaruc, J. & Onyaric, J. - Study on the microstructure of African wild silk cocoon shells and fibers International Journal of Biological Macromolecules, 50 (1): 63–68 access

2012 Teshome, A., Onyari, J., Raina, S., Kabaru, J. & Vollrath, F. - Mechanical and thermal degradation properties of silk from African wild silkmoths Journal of Applied Polymer Science, 127 (1): 289–297 access

2012 Chaudhury S., Dicko C., Vollrath F. & Carr A. - Determining chemical differences between torn and normal rotator cuff tendons using forier transfrom infared spectroscopy Orthopaedic Proceedings, 94-B access

2012 Chaudhury, S., Holland, C., Porter, D., Vollrath, F. & A J Carr. - Are they tough enough? A comparison of how the shear mechanical properties of rotator cuff repair patches match normal and torn cuff tendons Orthopaedic Proceedings, 94-B access

2012 Guan, J., Porter, D. & Vollrath, F. - Silks cope with stress by tuning their mechanical properties under load. Polymer, 53 (13): 2717–2726 access

2012 Chen, F., Porter, D. & Vollrath, F. - Silk Cocoon: Multilayer structure and mechanical properties (Bombyx mori). Acta Biomaterialia, 8 (7): 2620-2627 access

2012 Chen, F. J., Porter, D. & Vollrath, F. - Morphology and structure of silkworm cocoons. Materials Science & Engineering C-Materials for Biological Applications, 32 (4): 772-778 access

2012 Kronenberger, K., Moore, P.G., Halcrow, K. & Vollrath, F. - Spinning a marine silk for the purpose of tube-building. Journal Of Crustacean Biology, 32 (2): 191-201 access

2012 Holland, C., Vollrath, F. & Dicko, I.C. - Distinct structural and optical regimes in natural silk spinning. Biopolymers, 97 (6): 368-73 access

2012 Holland, C., Urbach, J. & Blair, D. - Direct Visualization of Shear Dependent Silk Fibrillogenesis. Soft Matter, 8 (9): 2590-4 access

2012 Holland, C., Porter, D. & Vollrath, F. - Comparing the rheology of mulberry and ‘wild’ silkworm spinning dopes. Biopolymers, 97 (6): 362-7 access

2012 Holland, C., Vollrath, F., Ryan, A.J. & Mykhaylyk, O.O. - Silk and Synthetic Polymers; Reconciling 100 Degrees of Separation. Advanced Materials, 24 (1): 105-109 access

2012 Greving, I., Cai, F., Vollrath, F. & Schniepp, H. - Shear-induced self-assembly of native silk proteins into fibrils studied by atomic force microscopy. Biomacromolecules, 13 (3): 676-682 access

2012 Huang, W., Begum, R., Barber, T., Ibba, V., Tee, N.C., Hussain, M., Arastoo, M., Yang, Q., Robson, L.G., Lesage, S., Gheysens, T., Skaer, N.J., Knight, D.P. & Priestley, J.V. - Regenerative potential of silk conduits in repair of peripheral nerve injury in adult rats. Biomaterials, 33 (1): 59-71 access


2011 Boulet-Audet, M. & Holland, C. - The African crested rat poisonous hairs studied by attenuated total reflection infrared spectroscopy. Spectroscopy Europe, 3 (6): 10-12 access

2011 Vollrath, F. - Super-materials made by silkworms. Oxford Impacts, access

2011 Turner, J., Vollrath, F. & Hesselberg, T. - Wind speed affects prey-catching behaviour in an orb web spider. Naturwissenschaften, 98 (12): 1063-1067 access

2011 Kronenberger, K., Dicko, C. & Vollrath, F. - A novel marine silk. Naturwissenschaften, 99 (1): 3-10 access

2011 Guan, J., Vollrath, F. & Porter, D. - Two Mechanisms for Supercontraction in Nephila Spider Dragline Silk. Biomacromolecules, 12 (11): 4030–4035 access

2011 Chaudhury, S., Dicko, C., Burgess, M., Vollrath, F. & Carr, A.J. - Fourier transform infrared spectroscopic analysis of normal and torn rotator-cuff tendons. The Journal of Bone and Joint Surgery, 93 (3): 370-377 access

2011 Chaudhury, S., Holland, C., Thompson, M., Vollrath, F. & Carr, A. - Tensile and Shear Mechanical Properties of Rotator Cuff Repair Patches. Journal of Shoulder and Elbow Surgery, 21 (9): 1168–1176 access

2011 Vollrath, F., Porter, D. & Holland, C. - There are many more lessons to be learned from spider silk. Soft Matter, 7 (20): 9595-9600 access

2011 Chen, F. & Hodgkinson, J. - "Impact damage" in Encyclopaedia of Composites, Wiley Publishing, in press

2011 Kingdon, J., Agwanda, B., Kinnaird, M., O’Brien, T., Holland, C., Gheysens. T., Boulet-Audet, M. & Vollrath, F. - A Poisonous Surprise Under The Coat Of The African Crested Rat. Philosophical Transactions of the Royal Society B, 279 (1729): 675-80 access

2011 Gheysens, T., Collins, A., Raina, S., Vollrath, F. & Knight, D.P. - Demineralization Enables Reeling of Wild Silkmoth Cocoons. Biomacromolecules, 12 (6): 2257-2266 access

2011 Teshome, A., Raina, S.K., Vollrath, F., Kabaru, J.M., Onyari, J. & Nguku, E.K. - Study on Weight Loss and Moisture Regain of Silk Cocoon Shells and Degummed Fibers from African Wild Silkmoths. Journal of Entomology, 8 (5): 450-458 access

2011 Chaudhury, S., Holland, C., Porter, D., Tirlapur, T., Vollrath, F. & Carr, A. - Torn Human Rotator Cuff Tendons Have Reduced Collagen Thermal Properties on Differential Scanning Calorimetry. Journal of Orthopedic Research, 29 (12): 1938-1943 access

2011 Chaudhury, S., Holland, C., Vollrath, F. & Carr, A. - Comparing normal and torn rotator cuff tendons using dynamic shear analysis. Journal of Bone and Joint Surgery, 93-B (7): 942-948 access

2011 Seydel, T., Knoll, W., Greving, I., Dicko, C., Koza, M.M., Krasnov, I. & ller, M. - Increased molecular mobility in humid silk fibers under tensile stress. Physical Review E, 83 (1): 016104 access

2011 Vollrath F. - "Spider Silks" in Encyclopedia of Nanotechnology, Spinger Verlag, Heidelberg/New York (in press) Reserve your copy now

2011 Boulet-Audet, M., Vollrath, F. & Holland, C. - Rheo-Attenuated Total Reflectance Infrared Spectroscopy: a new tool to study biopolymers. Physical Chemistry Chemical Physics, 13 (9): 3979-3984 access

2011 Fu, C., Porter, D., Chen, X., Vollrath, F. & Shao, Z. - Understanding the Mechanical Properties of Antheraea Pernyi Silk—From Primary Structure to Condensed Structure of the Protein. Advanced Functional Materials, 21 (4): 729-737 access


2010 Guan, J., Porter, D., Tian, K., Shao, Z.Z. & Chen, X. - Morphology and mechanical properties of soy protein scaffolds made by directional freezing. Journal of Applied Polymer Science, 118 (3): 1658–1665 access

2010 Yin, Y., Chen, E., Porter, D. & Shao, Z. - Enhancing the Toughness of Regenerated Silk Fibroin Film through Uniaxial Extension. Biomacromolecules, 11 (11): 2890-2895 access

2010 Dicko C., Porter D. & Vollrath F. - The relevance of silks to amyloids. pp51 -70 in Functional Amyloid Aggregation Eds: Stefania Rigacci and Monica Bucciantini. Research Signpost. ISBN: 978-81-308-0425-5 in press.

2010 Foreman, J.P., Porter, D., Behzadi, S., Curtis, P.T. & Jones, F.R. - Predicting the thermomechanical properties of an epoxy resin blend as a function of temperature and strain rate. Composites Part A: Applied Science and Manufacturing, 41 (9): 1072-1076 access

2010 Greving, I., Dicko, C., Terry, A.E., Callow, P. & Vollrath, F. - Small angle neutron scattering of native and reconstituted silk fibroin. Softmatter, 6 (18): 4389-4395 access

2010 Chen, F., Porter, D. & Vollrath, F. - A nonwoven composite model based on silkworm cocoon (Bombyx mori). Journal of Materials Science and Engineering, 4 (9): 28-33 Email for access

2010 Vollrath, F. - Silk evolution untangled Nature 466: 319 access

2010 Chen, F., Porter, D. & Vollrath. - Silkworm cocoons inspire models for random fiber and particulate composites. Physical Review E 82 (4): 0419111-0419116 access

2010  Hakimi, O., Gheysens, T., Vollrath, F., Grahn, M., Knight, D. & Vadgama, P. - Modulation of cell growth on exposure to silkworm and spider silk fibers. Journal of Biomedical Materials Research Part A, 92A (4): 1366–1372 access


2009 Fu, C., Porter, D. & Shao, Z. - Moisture Effects on Antheraea pernyi Silk’s Mechanical Property. Macromolecules, 42 (20): 7877-7880 access

2009 Collins, A., Skaer, N., Gheysens, T., Knight, D., Bertram, C., Roach, H., Oreffo, H., Von-Aulock, S., Baris, T., Skinner, J. & Mann, S. - Bone-like Resorbable Silk-based Scaffolds for Load-bearing Osteoregenerative Applications. Advanced Materials, 21 (1): 75-78 access

2009 Clifton, L.A., Neylon C., Terry A.E., Dicko C. & Diddens I. - Characterising biomaterial complexity. Materials Today, 12 (7-8): 86-91 access

2009 Vollrath, F., Chen, F. & Porter, D. - Silks and their composites. pp1355-1365 in Transactions of the 14th Fundamental Research Symposium on advances in pulp and paper research, Oxford 2009. ISBN 978 0 9545272 7 3 Email for access

2009 Fu, C., Shao, Z. & Vollrath, F. - Animal silks: their structures, properties and artificial production. Chemical Communications, (43): 6515-6529 access

2009 Porter, D. & Gould, P.J. - Predictive nonlinear constitutive relations in polymers through loss history. International Journal of Solids and Structures, 46 (9): 1981-199 access

2009 Vollrath, F., Porter, D. & Dicko, C. - The structure of silk. pp146-198 in Handbook of Textile Fibre Structure Vol.2 Eds S.J. Eichhorn, J.W.S. Hearlem, M. Jaffe & T. Kikutani. Woodhead Publishing access

2009 Vollrath, F. & Porter, D. - Silks as Ancient Models for Modern Polymers. Polymer, 50 (24): 5623-32 access

2009 Mo, C., Holland, C., Porter, D., Shao, Z. & Vollrath, F. - Concentration state dependence of the rheological and structural properties of reconstituted silk. Biomacromolecules, 10 (10): 2724-8 access

2009 Porter, D., Vollrath, F., Tian, K., Chen, X. & Shao, Z. - A kinetic model for thermal degradation in polymers with specific application to proteins. Polymer, 50 (7): 1814-1818 access

2009 Foreman, J.P., Behzadi, S., Tsampas, S.A., Porter, D., Curtis, P.T. & Jones, F.R. - Rate dependent multiscale modelling of fibre
reinforced composites. Plastics, Rubber and Composites, 38 (2-4): 67-71 access

2009 Porter, D. & Gould, P. J. - Predictive nonlinear constitutive relations in polymers through loss history. International Journal of Solids and Structures, 46 (9): 1981-1993 access

2009 Porter, D. & Vollrath, F. - Silk as a Biomimetic Ideal for Structural Polymers. Adv. Mater., 21 (4): 487-492 access


2008 Dicko, C., Hicks, M.R., Dafforn, T.R., Vollrath, F., Rodger, A. & Hoffmann S.V. - Breaking the 200 nm Limit for Routine Flow Linear Dichroism Measurements Using UV Synchrotron Radiation. Biophysical Journal, 95 (12): 5974-5977 access

2008 Cheng, C., Shao, Z. & Vollrath, F. - Silk Fibroin regulating the crystallization process of calcium carbonate. Adv. Funct. Mater., 18 (15): 2172-2179 access

2008 Dicko, C., Porter, D., Bond, J., Kenney, J. & Vollrath, F. - Structural Disorder in Silk Proteins Reveals the Emergence of Elastomericity Biomacromolecules, 9 (1): 216-221 access

2008 Liu, Y., Sponner, A., Porter, D. & Vollrath, F. - Proline and processing of spider silks. Biomacromolecules, 9 (1): 116-121  access

2008 Liu, Y., Shao, Z. & Vollrath, F. - Elasticity of Spider Silks Biomacromolecules, 9 (7): 1782–1786 access

2008 Porter, D. & Vollrath, F. - The role of kinetics of water and amide bonding in protein stability. Soft Matter, 4 (2): 328-336 access

2008 Brookes, V., Young, R. & Vollrath F. - Deformation micromechanics of spider silk Journal of Materials Science, 43:3728 access

2008 Holland, C. & Vollrath F. - Biomimetic principles of spider silk for high performance fibres. Chapter 7 in Biologically Inspired Textiles. Eds Ellison, M.S., Abbott, A.G., Woodhead Publishing


2007 Swinerd, V.M., Collins, A.M., Skaer, N.J.V., Gheysens, T. & Mann, S. - Silk inverse opals from template-directed b-sheet transformation of regenerated silk fibroin. Softmatter, 3 (11): 1377-1380. access

2007 Vollrath, F. & Selden, P. - The Role of Behavior in the Evolution of Spiders, Silks, and Webs. Annual Review of Ecology, Evolution, and Systematics, 38: 819-46 access

2007 Chen, X., Shao, Z., Knight, D.P. & Vollrath, F. - Conformation Transition Kinetics of Bombyx mori Silk Protein. Proteins: Structure, Function, and Bioinformatics, 68 (1): 223–231 access

2007 Porter, D. & Vollrath, F. - Nanoscale toughness of spider silk. Nanotoday, 2 (3): 6 access

2007 Emile, O., Floch, A.L. & Vollrath, F. - Time-resolved torsional relaxation of spider draglines by an optical technique. Physcial Review Letters, 98 (16): 167402-(1-4) access

2007 Holland, C., Terry, A., Porter, D. & Vollrath, F. - Natural and Unnatural Silks. Polymer, 48 (12): 3388-3392 access

2007 Hakimi, O., Knight, D., Vollrath, F. & Vadgama, P. - Spider and mulberry silkworm silks as compatible biomaterials. Composites Part B: Engineering, 38 (3): 324-337 access

2007 Huang, W.L., Begum, R., Yang, Q., Gheysens, T., Skaer, N., King, V., Knight, D. & Priestley, J.V. - Regenerative Potential of Silk Conduits Following Peripheral Nerve Injury. Tissue Engineering, 13 (7): 1633-1778 access


2006 Grip, S., Rising, A., Nimmervoll, H., Storckenfeldt, E., Mcqueen-mason, S.J., Pouchkina-stantcheva, N., Vollrath, F., Engström, W. & Fernandez-arias, A. - Transient Expression of a Major Ampullate Spidroin 1s Gene Fragment from Euprosthenops sp. in Mammalian Cells, Cancer Genomics & Proteomics, 3: 84-88 access

2006 Dicko, C., Kenney, J. & Vollrath, F. - Beta-Silks: Enhancing and controlling aggregation. Fibrous Proteins: Amyloids, Prions and Beta- proteins; Advances in Protein Chemistry, 73: 17-53 access

2006 Holland, C., Terry, A., Porter, D. & Vollrath, F. - Comparing the rheology of native spider and silkworm dope. Nature Materials, 5 (11): 870-874 access

2006 Chen, X., Shao, Z.Z. & Vollrath, F. - Spinning processes for spider silk. Soft Matter, 2 (6): 448–451 access

2006 Emile, O. Floch, A.L. & Vollrath, F. - Shape memory in spider draglines. Nature, 440 (7084): 621 access

2006 Vollrath, F. - Spider Silk: Thousands of Nano-Filaments and Dollops of Sticky Glue. Current Biology, 16 (21): R925-R927 access

2006 Vollrath, F. & Porter, D. - Spider silk as archetypal protein elastomer. Soft Matter, 2 (5): 377-385 access

2006 Vollrath, F. & Porter, D. - Spider silk as a model biomaterial. Applied Physics A: Materials Science & Processing, 82 (2): 205-212 access


2005 Sapede, D., Seydel, T., Forsyth, T., Koza, M.M., Schweins, R., Vollrath, F. & Riekel,C. - Nanofibrillar structure and molecular mobility in spider dragline silk. Macromolecules, 38 (20): 8447-8453 access

2005 Liu, Y., Shao, Z.Z. & Vollrath, F. - Relationships between supercontraction and the mechanical properties of spider silk. Nature Materials, 4 (12): 901-905 access

2005 Liu, Y., Shao, Z. & Vollrath, F. - Extended wet-spinning can modify spider silk properties. Chemical Communications, (19): 2489-91 access

2005 Dicko, C., Knight, D., Kenney, J. & Vollrath, F. - Conformational polymorphism, stability and aggregation in spider dragline silk proteins. International Journal of Biological Macromolecules, 36 (4): 215-224 access

2005 Sponner, A., Vater, W., Rommerskirch, W., Vollrath, F., Unger, E., Grosse, F. & Weisshart, K. - The conserved C-termini contribute to the properties of spider silk fibroins. Biochem. Biophys. Res. Commun., 338 (2): 897-902 access

2005 Sponner, A., Unger, E., Grosse, F. & Weisshart, K. - Differential polymerization of the two main protein components of dragline silk during fibre spinning. Nat. Mat., 4 (10): 772-5 access

2005 Sponner, A., Schlott, B., Vollrath, F., Unger, E., Grosse, F. & Weisshart, K. - Characterization of the Protein Components of Nephila clavipes dragline silk. Biochemistry, 44 (12): 4727-4736 access

2005 Peng, X., Shao, Z., Chen, X., Knight, D.P., Wu, P. & Vollrath, F. - Further Investigation on Potassium-Induced Conformation Transition of Nephila Spidroin Film with Two-Dimensional Infrared Correlation Spectroscopy. Biomacromolecules, 6 (1): 302-306 access

2005 Porter, D., Vollrath, F. & Shao, Z. - Predicting the mechanical properties of spider silk as a model nanostructured polymer. Europ. Physical J. E, 16 (2): 199-206 access

2005 Vollrath, F. - Spiders' Webs. Current Biology, 15 (10): R364-R365 access

2005 Yang, Y., Chen, X., Zhou, P., Shao, Z., Porter, D., Knight, D.P. & Vollrath, F. - Toughness of spider silk at high and low temperatures. Advanced Materials, 17 (1): 83-88 access

2005 Vollrath, F, Sponner A, A Synthetic natural silks. - ‘The route to synthetic silks’ 245-270, in ‘Biodegradable and sustainable fibres’ Editor R.Blackburn, Woodhead publishing Limited, Cambridge, UK (ISBN-13: 978-1-85573-916-1, ISBN-10: 1-85573-916-X )


2004 Sponner, A., Unger, E., Grosse, F. & Weisshart, K. - Conserved C-termini of spidroins are secreted by the major ampullate glands and retained in the silk thread. Biomacromol., 5 (3): 840-5 access

2004 Vollrath, F. & Knight, D.P. - Biology and technology of silk production (in press) in (eds A. Steinbüchel and Y. Doi) Biotechnology of biopolymers : from synthesis to patents. Wiley-VCH, Heidelberg & New York (reprinted article in Handbook of Biopolymers 2003)

2004 Terry, A., Knight, D.P., Porter, D. & Vollrath, F. - pH induced changes in the rheology of silk fibroin solution from the middle division of Bombyx mori Silkworm. Biomacromolecules, 5 (3): 268-277 access

2004 Huemmerich, D., Scheibel, T., Vollrath, F., Cohen, S., Gat, U. & Ittah S. - Novel Assembly Properties of Recombinant Spider Dragline Silk Proteins. Current Biology, 14 (22): 2070-2074 access

2004 Riekel, C., Roessle, M., Sapede, D. & Vollrath, F. - Influence of CO2 on the micro-structural properties of spider dragline silk: X-ray microdiffraction results. Naturwissenschaften, 91 (1): 30-33 access

2004 Hronsky, M., van Beek, J.D., Williamson, P.T.F., Vollrath, F. & Meier, B.H. - NMR characterization of native liquid spider dragline silk from Nephila edulis. Biomacromol., 5 (3): 834-839 access

2004 Hesselberg, T. & Vollrath, F. - The Effects of Neurotoxins on Web-geometry and Web-building Behaviour in Araneus diadematus Cl. Physiology and Behaviour, 82 (2-3): 519-529 access

2004 Dicko, C., Knight, D., Kenney, J.M. & Vollrath, F. - Transition to a beta-sheet-rich Structure in Spidroin in Vitro:  The Effects of pH and Cations. Biochemistry, 43 (44): 14080-87 access

2004 Dicko, C., Kenney, J.M. & Vollrath, F. - Secondary Structures and Conformational Changes in Flagelliform, Cylindrical, Major, and Minor Ampullate Silk Proteins. Temperature and Concentration Effects. Biomacromolecules, 5 (6): 2105-2115 access

2004 Dicko, C., Knight, D., Kenney, J.M. & Vollrath, F. - Structural conformation of spidroin in solution: A synchrotron radiation circular dichroism study. Biomacromolecules, 5 (3): 758-767 access

2004 Dicko, C., Kenney, J.M. & Vollrath, F. - Spider silk-protein refolding is controlled by changing pH. Biomacromolecules, 5 (3): 704-710 access


2003 Zhou, L., Chen, X., Shao, S., Zhou, P., Knight, D.P. & Vollrath, F. - Copper in the silk formation process of Bombyx mori silkworm. FEBS Letters, 554 (3): 337-341 access

2003 Vollrath, F. & Knight, D. - Biology and technology of silk production. pp 25-46 in Handbook of Biopolymers. (eds. A Steinbüchel, S Fahnestock), Wiley-VCH, Heidelberg & New York

2003 Vollrath, F. & Knight, D. - The nature of some spiders' silks. pp 152-174 in (eds B Shrewry, A.Tatham,) Elastomeric Proteins, Cambridge University Press, Cambridge, UK

2003 Knight, D.P. & Vollrath, F. - Biological liquid crystal elastomers. pp 302-320 reprinted in (eds B Shrewry, A.Tatham,) Elastomeric Proteins, Cambridge University Press, Cambridge, UK, (reprinted article from Phil. Trans. Roy. Soc. 2002) access

2003 Knight, D.P. & Vollrath, F. - Spinning an elastic ribbon of spider silk. pp115-135 reprinted in (eds B Shrewry, A.Tatham,) Elastomeric Proteins, Cambridge University Press, Cambridge, UK, (reprinted article from Phil. Trans. Roy. Soc. 2002) access

2003 Chen, X., Zhou, L., Shao, S., Zhou, P., Knight, D.P. & Vollrath, F. - Conformation Transition of Silk Protein Membranes Monitored by Time-resolved FTIR Spectroscopy: Conformation Transition Behavior of Regenerated Silk Fibroin Membranes in Alcohol Solution at High Concentration. Acta Chimica Sinica, 61 (4): 625-629. abstract

2003 Shao, Z., Yang, Y. & Vollrath, F. - Structure and behaviour of regenerated spider silk. Macromolecules, 36 (4): 1157-1161 access


2002 Vollrath, F., Barth, P., Basedow, W., Engström, W. & List, H. - Local tolerance to spider silks and protein polymers. In Vivo, 16 (4): 229-234 abstract

2002 Vollrath, F. - Microstructure of spider silks. pp 93-106 in Sustainable Natural and Polymeric Composites - Science and Technology (eds H Lilholt et al.) Risø Internatl. Symp. Mat. Sci. Riso National Laboratories, Roskilde

2002 Van Beek, J.D., Hess, S., Vollrath, F. & Maier, B.H. - The molecular structure of spider dragline silk: folding and orientation of the protein backbone. Proc. Nat. Acad. Sci., 99 (16): 10266-10271 access

2002 Shao, Z. & Vollrath, F. - Surprising strength of silkworm silk. Nature, 418 (6899): 741 access

2002 Knight, D.P. & Vollrath, F. - Spinning an elastic ribbon of spider silk. Phil. Trans. Roy. Soc., 357 (1418): 219-228 access

2002 Knight, D.P. & Vollrath, F. - Biological liquid crystal elastomers. Phil. Trans. Roy. Soc., 357 (1418): 155-163 access

2002 Kenney, J., Knight, D.P., Wise, M. & Vollrath, F. - Amyloidogenic nature of spider silk. Eur. J. Biochem., 269 (16): 4159–4163 access

2002 Chen, X., Shao, Z., Knight, D.P. & Vollrath, F. - Conformation Transition of Silk Protein Membranes Monitored by Time-resolved FTIR Spectroscopy: Effect of Alkali Metal Ions on Nephila Spidroin Membrane. Acta Chimica Sinica, 60 (12): 2203-2208 abstract

2002 Chen, X., Knight, D.P. & Vollrath, F. - Rheological characterization of Nephila spidroin solution. Biomacromol., 3 (4): 644-648 access

2002 Chen, X., Knight, D.P., Shao, Z. & Vollrath, F. - Conformation Transition in Silk Protein Films Monitored by Time-Resolved Fourier Transform Infrared Spectroscopy: Effect of Potassium Ions on Nephila Spidroin Films. Biochemistry, 41 (5): 14944-14950 access


2001 Vollrath, F., Madsen, B. & Shao, Z. - The effect of spinning conditions on the mechanics of a spider's dragline silk. Proc. Roy. Soc., 268 (1483): 2339-2346 access

2001 Vollrath, F. & Knight, D.P. - Liquid crystalline spinning of spider silk. Nature, 410 (6828): 541-548 access

2001 Riekel, C. & Vollrath, F. - Spider silk fibre extrusion: combined wide- and small-angle X-ray microdiffraction experiments. Int. J. Biol. Macromol., 29 (3): 203-210 access

2001 Knight, D.P. & Vollrath, F. - Comparison of liquid crystal spinning of Selachian egg case ply sheets and orb web spider dragline filaments. Biomacromol., 2 (2): 323-334 access

2001 Knight, D.P. & Vollrath, F. - Changes in element composition along the spinning duct in a Nephia spider. Naturwissenschaften, 88 (4): 179-182 access

2001 Chen, X., Knight, D.P., Shao, Z. & Vollrath, F. - Regenerated Bombyx silk solutions studied with rheometry and FTIR. Polymer, 42 (25): 9969-9974 access


2000 Vollrath, F. - Strength and structure of spiders' silks. Rev. Molec. Biotech., 74 (2): 67-83 access

2000 Vollrath, F., Nørgaard, T. & Krieger, M. - Radius orientation in the cross spider Araneus diadematus. Pp 107-116 in European Arachnology (eds S Toft & N Scharff) University Press, Aarhus

2000 Vollrath, F. - Coevolution of behaviour and material in the spider's web. pp 315-329 in Biomechanics in Animal Behaviour (ed. P Domenici, RW Blake). Bios, Oxford

2000 Uhl, G. & Vollrath, F. - Extreme body size variability in the golden silk spider (Nephila edulis) does not extend to genitalia. J. Zoology, 251 (1): 7-14 access

2000 Riekel, C., Madsen, B., Knight, D.P. & Vollrath, F. - X-ray Diffraction on Spider Silk during Controlled Extrusion under a Synchrotron Radiation X-ray Beam. Biol. Macromol., 1 (4): 622-626 access

2000 Madsen, B. & Vollrath, F. - Mechanics and morphology of silk drawn from anaesthetised spiders. Naturwiss., 87 (3): 148-153 access

2000 Krink, T. & Vollrath, F. - Optimal Area Use in Orb Webs of the Spider Araneus diadematus. Naturwiss., 87 (2): 90-93. access

2000 Knight, D.P., Knight, M.M. & Vollrath, F. - Beta transition and stress-induced phase separation in the spinning of spider dragline silk. Int. J. Biol. Macromol., 27 (3): 205-210 access

2000 Kenney, J., Knight, D.P., Dicko, C. & Vollrath, F. - Linear and circular dichroism can help us to understand the molecular nature of spider silk. pp 123-126. Proceedings of the19th European Arachnology Conference access


1990s <back to top>

1999 Vollrath, F. & Knight, D.P. - Structure and function of the silk production pathway in the spider Nephila edulis. Int. J. Biol. Macromol., 24 (2-3): 243-249 access

1999 Vollrath, F. - Biology of spider silk. Int. J. Biol. Macromol., 24 (2-3): 81-88 access

1999 van Beek, J.D., Kümmerlen, J., Vollrath, F. & Meier, B.H. - Supercontracted spider dragline silk: a solid state NMR study of the local structure. Int. J. Biol. Macromol., 24 (2-3): 173-178 access

1999 Sirichaisit, J., Young, R.J. & Vollrath, F. - Molecular deformation in spider dragline silk subjected to stress. Polymer, 41 (3): 1223-1227. access

1999 Shao, Z., Young, R.J. & Vollrath, F. - The effects of solvents on spider silk studied by mechanical testing and single-fibre Raman spectroscopy. Int. J. Biol. Macromol., 24 (2-3): 295-300 access

1999 Shao, Z. & Vollrath, F. - The effect of solvents on the contraction and mechanical properties of spider silk. Polymer, 40 (7): 1799-1806 access

1999 Shao, Z., Vollrath, F., Sirichaisit, J. & Young, R.J. - Analysis of spider silk in native and supercontracted states using Raman spectroscopy. Polymer, 40 (10): 2493-2500 access

1999 Shao, Z., Hu, X.W., Frische, S. & Vollrath, F. - Heterogeneous morphology in Nepila edulis spider silk and its significance for mechanical properties. Polymer, 40 (16): 4709-4711 access

1999 Riekel, C., Müller, M. & Vollrath, F. - In-situ X-ray diffraction studies during forced silking spider silk. Macromolecules, 32 (14): 4464-4466 access

1999 Madsen, B., Shao, Z. & Vollrath, F. - Variability in the mechanical properties of spider silks on three levels: interspecific, intraspecific and intraindividual. Int. J. Biol. Macromol., 24 (2-3): 301-306 access

1999 Krink, T. & Vollrath, F. - A virtual robot to model the use of regenerated legs in a web-building spider. Anim. Behav., 57 (1): 223-241 access

1999 Krink, T. & Vollrath, F. - Spatial Orientation of virtual Spider Robots by Object-oriented Rule-based Simulations. IEEE Intelligent Systems, 14 (5): 77-84 access

1999 Knight, D.P. & Vollrath, F. - Liquid crystals and flow elongation in a spider's silk production line. Proc. Roy. Soc., 266 (1418): 519-523 access

1999 Knight, D.P. & Vollrath, F. - Hexagonal columnar liquid crystals in the cells secreting spider silk. Tissue and Cell, 31 (6): 617-620 access

1999 Frische, S., Maunsbach, A. & Vollrath, F. - Elongate cavities and skin-core structure in Nephila spider silk observed by electron microscopy. J. Microscopy, 189 (1): 64-70 access


1998 Young, R.J., Sirichaisit, J., Shao, Z. & Vollrath, F. - Analysis of structure/property relationships in spider silk using Raman spectroscopy. pp 1-7 in The New Polymers: Science, Technology & Applications, Institute of Physics, Brighton

1998 Weissmann. M. & Vollrath, F. - The effect of leg loss on orb spider growth. Bull. Brit. Arachnol. Soc., 11 (3): 92-94 BAS members access

1998 Vollrath, F. - Sex-linked differences in the pattern of growth in Nephila clavipes. pp 161-66 in: Proc. 17th Europ. Coll. Arachnol. (Edinburgh) access

1998 Vollrath, F. - Dwarf males. Trend. Ecol. Evolution, 13 (4): 159-163 access

1998 Vollrath, F., Hu, X.W. & Knight, D.P. - Silk production in a spider involves acid bath treatment. Proc. R. Soc. B., 265 (1398): 817-820 access

1998 Uhl, G. & Vollrath, F. - Little evidence for size dependant sexual cannibalism in two Nephila species. Zoology, 101: 101-106

1998 Uhl, G. & Vollrath, F. - Genital morphology of Nephila edulis: implications for sperm competition in spiders. Can. J. Zool., 76 (1): 39-47 access

1998 Schneider, J. & Vollrath, F. - The effect of prey type on the geometry of the capture web of Araneus diadematus. Naturwissenschaften, 85 (8): 391-394 access

1998 Maynes, E., Mann, S. & Vollrath, F. - Fabrication of magnetic spider silk and other silk-fiber composites using inorganic nanoparticles. Advanced Materials, 10 (10): 801-805 access

1998 Krink, T. & Vollrath, F. - Emergent properties in the behaviour of a virtual spider robot. Proc. R. Soc. Lond. B, 265 (1410): 2051-2055 access


1997 Vollrath, F. & Samu, F. - The effect of starvation on the web geometry of an orb weaving spider. Bull. Brit. Arachnol. Soc., 10 (8): 295-298 BAS members access

1997 Vollrath, F. & Krink, T. - En edderkops gang gennem det evolutionaere landskab. Naturens Verden, 1997 (11/12): 440-453

1997 Vollrath, F. & Parker, G.A. - Giant female or dwarf male spiders? Nature, 385 (6618): 688 access

1997 Vollrath, F., Downes, M. & Krakow, S. - Design variability in web-geometry of an orb-weaving spider. Physiol. Behav., 62 (4): 735-743 access

1997 Shao, Z. & Vollrath, F. - The mechanics of conditioned silks. pp 262-265 in Deformation, Yield and Fracture of Polymers. The Institute of Materials, London

1997 Krink, T. & Vollrath, F. - Analysing spider web-building behaviour with rule-based simulations and genetic algorithms. J. theor. Biol., 185 (3): 321-331 access

1997 Hutchinson, J.M., McNamara, J.M., Houston, A.I. & Vollrath, F. - Dyar's rule and the investment principle: Optimal moulting strategies if feeding rate is size-dependent and growth is discontinous. Phil. Trans. Roy. Soc. B, 352 (1349): 113-138 access


1996 Vollrath, F. - Edderkoppespind - et kemisk og biomechanisk viduer. Naturens Verden, 10: 371-376

1996 Abraham, J., Medenwaldt, R., Uggerhoj, U., Guttmann, P., Hjort, T., Jensenius, J., Vorup-Jensen, T., Vollrath, F., Søgaard, E., Møller, T. - X-ray Microscopy in Aarhus. X-ray Microscopy and Spectromiscroscopy - XRM 96 (eds. J. Thieme et al.) Springer Verlag, Heidelberg

1996 Vollrath, F. & Köhler, T. - Mecahnics of silk produced by loaded spiders. Proc. Roy. Soc., 263 (1369): 387-391 access

1996 Vollrath, F., Holtet, T., Thøgersen, H.C. & Frische, S. - Structural organization of spider silk. Proc. Roy. Soc., 263 (1367): 147-151 access

1996 Kümmerlen, J., van Beek, J.D., Vollrath, F. & Meier, B.H. - The local structure in spider dragline silk investigated by two-dimensional spin-diffusion nuclear magnetic resonance. Macromolecules, 29 (8): 2920-2928 access


1995 Vollrath, F. & Edmonds, D. - Elastic properties of spider's capture silk. Naturwissenschaften, 82 (8): 379-380 access

1995 Vollrath, F. & Milinski, M. - Fragrant genes help Damenwahl. Trends in Ecology and Evolution, 10 (8): 307-308 access

1995 Downes, M. & Vollrath, F. - Foxes, hedgehogs and modern natural history. Flora og Fauna, 100: 123-126

1995 Zschokke, S. & Vollrath, F. - Web construction patterns in a range of orb weaving spiders. Europ. J. Entomol., 92 (3): 523-541 access

1995 Zschokke, S. & Vollrath, F. - Unfreezing the behaviour of web spiders. Behav. Physiol., 58 (6): 1167-1173 access

1995 Vollrath, F. - Lyriform organs on regenerated legs of the garden cross spider. Bull. Br. Arachnol. Soc., 10 (3): 115-118

1995 Lin, L., Edmonds D. & Vollrath F. - Structural engineering of a spider's web. Nature, 373 (6510): 146-148 access

1995 Köhler, T. & Vollrath, F. - Thread biomechanics in the two orb weaving spiders Araneus diadematus (Araneae, Araneidae) and Uloborus walckenaerius (Araneae, Uloboridae). J. exp. Zool., 271 (1): 1-17 access

1995 Fels, D., ap Rhisisart, A. & Vollrath, F. - The selfish crouton. Behavior, 132 (1): 49-55 access


1994 Vollrath, F. - General Properties of spider silk. pp 17-28 in. Silks Polymers: Materials Science and Biotechnology. (eds: Kaplan, D.L., Adams, W.W., Viney, C. & Farmer, B.L.) ACS Books, Washington

1994 ap Rhisiart, A. & Vollrath, F. - Design features of a spider's orb web. Behav. Ecol., 5 (3): 280-287 access

1994 Vollrath, F. - To catch a fly. Science and Public Affairs, 94: 20-24


1993 Vollrath, F. - Spinnenseide - Superwerkstoff der Natur pp.1-18 in : Kunststoffe im Automobilbau. VDI Verlag, Düsseldorf

1993 Vollrath, F. - Spidersilk - a natural polymer, and more. pp 1-16 in: Plastics in Engineering L. Vollrath ed., VDI Verlag, Düsseldorf


1992 Vollrath, F. & Parker, G.A. - Sexual dimorphism and distorted sex ratios in spiders. Nature, 360 (6400): 156-159 access

1992 Vollrath, F. - Spider webs and silks. Scientific American, March, 266:3 46-52 access

1992 Vollrath, F. - Die Seiden und Netze von Spinnen. Spektrum der Wissenschaft, May, pp 82-89

1992 Vollrath, F. - Analysis and interpretation of orb spider exploration and web-building behaviour. Adv. Study Behav., 21: 147-199 access

1992 Samu, F., Matthews, G., Lake, D. & Vollrath, F. - Spider webs are efficient collectors of agrochemical spray. Pesticide Sci., 36 (1): 47-51 access

1992 Samu, F. & Vollrath, F. - Spider orb webs as bioassays for pesticide side effects. Ent. Appl. Exp., 62 (2): 117-124 access

1992 Hurst, L. & Vollrath, F. - Sex ratio adjustment in solitary and social spiders. Trends Ecol. Evol., 7 (10): 326-327 access

1992 Gotts, N. & Vollrath, F. - Physical and theoretical features in the simulation of animal behaviour, e.g. the spider’s web. Cybernet. System. 23 (1): 41-65 access

1992 Edmonds, D. & Vollrath, F. - The contribution of atmospheric water vapour to the formation and efficiency of a spider's capture web. Proc. Roy. Soc. London, 248 (1322): 145-148 access

1992 Bonthrone, K.M., Vollrath, F., Hunter, B. & Sanders, J.K.M. - The elasticity of spider’s webs is due to water-induced mobility at a molecular level. Proc. Roy. Soc. London, 248 (1322): 141-144 access

1992 Vollrath, F. & Parker, G.A. - Sexual dimorphism and distorted sex ratios in spiders. Nature, 360 (6400): 156-159 access


1991 Vollrath, F. & Tillinghast, E. - Glycoprotein glue beneath a spider web's aqueous coat. Naturwissenschaften., 78 (12): 557-559 access

1991 Gotts, N. & Vollrath, F. - Artificial intelligence modelling of web-building in the garden cross spider. J. Theor. Biol., 152 (4): 485-511 access


1990 Vollrath, F. - Leg regeneration in web spiders and its implications for orb weaver phylogeny. Bull. Brit. Arachnol. Soc., 8 (6): 177-184

1990 Vollrath, F., Fairbrother, W.J., Williams, R.J.P., Tillinghast, E.K., Bernstein, D.T., Gallager, K.S. & Townley, M.A. - Compounds in the droplets of the orb spider's viscid spiral. Nature, 345 (6275): 526-528 access


1980s <back to top>


1989 Vollrath, F. & Edmonds, D. - Modulation of the mechanical properties of spider silk by coating with water. Nature, 340 (6231): 305-307 access


1988 Vollrath, F. - Spiral orientation of Araneus diadematus orb webs build during vertical rotation. J. Comp. Physiol. A, 162 (3): 413-419 access

1988 Vollrath, F. - Spider growth as an indicator of habitat quality. Bull. Br. Arachnol. Soc., 7 (7): 217-219


1987 Vollrath, F. - Kleptobiosis in spiders. pp 274-286 in Ecophysiology of Spiders. (ed. W. Nentwig), Springer Heidelberg access

1987 Vollrath, F. - Foraging, growth and reproductive success. pp 357-370 in Ecophysiology of Spiders. (ed. W Nentwig), Springer Heidelberg access

1987 Vollrath, F. - Untangling the spider's web. Trends Ecol. Evol., 3 (12): 331-335 access

1987 Vollrath, F. - Altered geometry of web in spiders with regenerated legs. Nature, 328 (6127): 247-248 access


1986 Vollrath, F. & Windsor, D. - Colony defence in a social and in a subsocial spider. pp 295-298, in: Proc. 9th Internatl. Arachnol. Congress. (WG Eberhard, YD Lubin, BC Robinson eds.), Smithsonian Press, Washington DC

1986 Vollrath, F. Houston A. - Previous experience and site tenacity in the orb spider Nephila clavipes. Oecologia, 70 (2):305-308 access

1986 Vollrath, F. - Gravity as orientation guide during web-construction in the orb spider Araneus diadematus. J. Comp. Physiol. A, 159 (2): 275-280 access

1986 Vollrath, F. - Eusociality and extraordinary sex ratios in the spider Anelosimus eximius. Behav. Ecol. Sociobiol., 18 (4): 283-287 access

1986 Vollrath, F. - Environment, reproduction and sex ratios in the social spider Anelosimus eximius. J. Arachnol., 14 (2): 267-281 access


1985 Vollrath, F. & Mohren, W. - Spiral geometry of the garden spider's orb web. Naturwissenschaften, 72 (12): 666-667 access

1985 Vollrath, F. - Web spider's dilemma: a risky move or site dependent growth. Oecologia, 68 (1): 69-72 access


1984 Vollrath, F. - Kleptoparasitism in Invertebrates. pp 61-94, in Producers and Scroungers: Strategies of Exploitation and Parasitism. (ed. C Barnard) Chrom Helm, acccess


1983 Vollrath, F. & Rohde-Arndt, D. - Prey capture and the distribution of food in the social spider Anelosimus eximius. Z. Tierpsychol., 61: 334-340 access

1983 Vollrath, F. - Relative and absolute growth in Nephila clavipes. pp 277-289 in Taxonomy, Biology and Ecology of Araneae. (ed. O Kraus), Verhandl. Naturwiss. Ver. Hamburg vol. 26 access


1982 Vollrath, F. - Colony foundation in a social spider. Z. Tierpsychol., 60:313-324 access


1981 Vollrath, F. - Energetic considerations of a spider parasite - spider host system. Rev. Arachnol. 3:37-44


1980 Vollrath, F. - Male body size and fitness in the web-building spider Nephila clavipes. Z. Tierpsychol., 53 (1): 61-78

1980 Vollrath, F. - Why are some spider males small? pp 165-169, in: Verh. 8th Internat.Arachnol. Congress, (ed. J Gruber), Egermann, Wien


1970s <back to top>


1979 Vollrath, F. - Vibrations: Their signal function for a spider kleptoparasite. Science, 205 (4411): 1149-1151 access

1979 Vollrath, F. - Behaviour of the kleptoparasitic spider Argyrodes elevatus. Animal Behav., 27-515-521 access


1978 Vollrath, F. - Behaviour of two kleptoparasitic species of the genus Argyrodes (Araneidae, Theridiidae). abstract in Symp. Zool. Soc. Lond., 42: 483 access

1978 Vollrath, F. - A close relationship between two spiders (Arachnida, Araneidae): Curimagua bayano synecious on a Diplura species. Psyche, 85: 347-353 access


1977 Vollrath, F. - Zur Ökologie Und Biologie Von Kleptoparasitischen Argyrodes Elavatus Und Synöken Argyrodes-Arten (Araneae, Theridiidae). Inaugural - Dissertation, Freiburg (Breisgau), Univ., Fak. für Biologie, Diss., (Nicht f. d. Austausch.) access


1976 Vollrath, F. - Konkurrenzvermeidung bei tropischen kleptoparasitischen Haubennetzspinnen der Gattung Argyrodes (Arachnida: Araneae: Theridiidae). Ent. Germ., 3 (1-2): 104-108 access

1976 Vollrath, F. - Competitive exclusion in tropical klepto parasitic spider species in the genus Argyrodes (Arachnida: Araneae: Theridiidae). Ent. Germ., 3 (1-2): 104-108 access

1976 Vollrath, F. & Delius J.D. - Vestibular projections to the thalamus of the pigeon. Brain, Behav. Evol., 13:58-68 access


1973 Delius, J.D., & Vollrath, F. - Rotation compensating reflexes independent of the labyrinth: neurosensory correlates in pigeons. J. Comp. Physiol. A. GSON 83:123-134 access



2005 Vollrath et al - Apparatus and method for forming materials Number 6858168 access

2006 Vollrath - Precursor silk feedstock for forming filaments Number US7041797 B2 access

Other Publications

2012 ???•??? <) ??:?????????
(The Selfish Gene by Richard Dawkins, translated by Fujia Chen)

2010 Hague, D. & Milnes, A. - From Bright Sparks to Brilliant Businesses, Case 12, Oxford Biomaterials, Suturox, Neurotex and Orthox, Fritz Vollrath. A & C Black Publishers Ltd. access


Recent News

December 2020

Two new studies have found that spiders can not only catch prey efficiently even when webs are severely distorted, but also build a normal web even when the structures to which it is anchored are continually moving (in press).

November 2020

This paper reviewed in Scientific American examines an interesting and potentially very important paradigm of morphological computing to study and test new ideas about the next generation of robots.

November 2019

Prof. Fritz Vollrath and colleagues from the Fudan University in China are widely covered in the news for their discovery of a means to produce fake Rhino horns using horse hair. Hopes are that this product may undermine the illegal market for rhino horn, and demistify the properties of rhino horn. View Here

October 2019

Scientists from the universities of Oxford, Shanghai and Beijing discovered how it can be that natural silks get stronger as they get colder. View Here

September 2019

Our latest Spin-out Spintex Engineering will use Aquamelt TM spinning technology to manufacture spider-type filaments


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Copyright 2007 OSG last updated 21 December, 2015