I attended Bolton School before reading Natural Sciences at Cambridge where I took the Natural Science Tripos in 1963 reading Biochemistry for Part II. After a short period in Switzerland I joined Philip Randle's group in the newly formed Biochemistry department at the University of Bristol. Here I took my PhD and became a Lecturer in Biochemistry. In 1975 I moved to Oxford where I was appointed Lecturer in Clinical Biochemistry and Tutor in Biochemistry for Magdalen and Worcester Colleges. I was made Reader in Clinical Biochemistry in 1990 and retired in 2002. Since retirement I have been a guest researcher at the Oxford Centre for Diabetes, Endocrinology and Metabolism

1960-63 Christ's College, Cambridge
1963-65 Clinique Universitaire Médicale, Lausanne
1965-69 MRC Research Assistant, University of Bristol
1969-71 Research Associate, University of Bristol
1971-75 Lecturer in Biochemistry, University of Bristol
1975-90 Lecturer in Clinical Biochemistry, University of Oxford
1975-90 Lecturer in Biochemistry, Worcester College, Oxford
1975-2002 Tutor in Biochemistry, Magdalen College, Oxford
1990-2002 Reader in Clinical Biochemistry, University of Oxford